To give a simple example suppose you have have a requirement to execute a system / os level command before or after a file is processed for some further operation then wht will you do ? Solution is that you specify the command to be executed in COMMAND LINE parameter.
One of the best examples is demonstrated by Michall where you have to zip a target file but don't want to use payloadzip bean then you can use command at the system level which can do the job.
Have a look here under "Run Operating System Command Before/After Message Processing"
Run Operating System Command Before/After Message Processing:
Command Line
The operating system command specified here is executed before or after successful file processing. The default value is an empty character string (no command).
When the operating system command is called, the file name currently being processed can be specified with the following placeholders:
%f (file name)
%F (absolute file name including path)
Check these blogs you have examples for OS command:
Run Operating System Command Before/After Message Processing -
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